Animals

Most animals that could be considered pathogens of plants are arthropods, in particular insects and mites. However, in the soil there is a whole world of pathogenic nematodes (microscopic worms), which form galls on roots. Some nematodes even form galls on leaves, perhaps dispersing through water splashing between plants.

Nematodes

Arthropods
Arachnids

Mites

Insects

Thrips

Bugs

Wasps

Beetles

Flies

I have left the vast majority of clades out of this tree. Those included are key groups of gall-formers.



Gall Midges

On flowers and fruits

Contarinia dipsacearum on Succisa pratensis.
Kiefferia pericarpiicola on Daucus carota.

On leaves and stems

Jaapiella veronicae on Veronica chamaedrys.
Jaapiella kiefferiana on Chamaenerion angustifolium.
Rabdophaga salicis on Salix sp.

Most midge galls consist of rolled-/curled-up leaves that enclose the larvae in a small chamber. Some form fully enclosed galls inside stems (right). The larvae are generally bright to pale orange.



Mites

Note I am not including non-galling mites such as Red Spider Mite here. These are also diverse and apparently at least somewhat host-specific.

Aceria thomasi on Thymus praecox.
Aceria anceps (I think...) on Veronica chamaedrys.
Eriophyes tiliae on Tilia sp.

Gall mites often cause the leaves of a plant to turn in on themselves and grow more hair than normal. On trees there are many species that form more classic ‘gall’-looking structures, also on the leaves (right).



Wasps

Neuroterus querc­us­bacca­rum on Quercus sp.
Andricus foecundatrix on Quercus sp.
Diplolepis spinosissimae on Rosa spinosissima.

Gall wasps are extremely diverse. Some wasp galls can be host to many different species, including the initial galler, commensals, parasitoids, and hyperparasitoids.



Aphids

Tetraneura ulmi on Ulmus sp.
Brachycolus cerastii on Cerastium sp.

Note I am only including galling species here.



Other animals

Nematodes

Subanguina graminophila on Agrostis stolonifera.
A juvenile of Subanguina graminophila under the microscope.

Most plant-parasitic nematodes form galls on the roots of plants, so you are unlikely to encounter them without searching. However, some form galls on leaves, such as Subanguina graminophila (pictured).


Thrips

Thrips are often host-specific and can cause gall-like symptoms.